Contents:
- Sexual Propagation
- Asexual Propagation
- Grafting Techniques
- Whip/Cleft Grafting
- T-Budding
- Chip Budding
Plants have two modes of propagation: sexual
and asexual
Sexual Propagation describes
when male genes (pollen grain) unite with female genes (ovules)
to form seed. There are two ways that this occurs:
- Self-pollination: when a plant's pollen fertilizes its own ovules;
the offspring are similar to parent.
- Cross-pollination: when a plant's pollen fertilizes ovules on
another plant - new gene combinations, offspring are genetically
different from either parent - unreliable, never know what you
will get: offspring can produce delicious new varieties of fruit
or be totally unproductive.
Pollen grain are transferred from the male flower part (stamen)
to female flower part (stigma), which leads to pollination. The
pollen grain then grows a tubule down through the stigma to the
ovary, and sends male genes into ovary to unite with female genes
in ovules, fertilizing it. The ovary then develops into a fruit,
with the ovules becoming individual seeds. Mature fruit falls to
ground, and seeds overwinter on or in the soil. In spring, the seeds
germinate and seedlings appear.
Modes of Pollination:
- Natural: Pollen grains are transported by insects, birds,
rain, and wind.
- Manual/artificial: Orchardists transfer pollen from one
flower to another. Offspring usually resemble parent plants, but
this may result in sterile hybrid plants.
Seed Stratification – procedures used to break seed dormancy.
- Most seeds germinate in response to cold soil temperature. To
simulate overwintering store seeds for 2-4 months in moist sand
or peat at near-freezing temperature (33-44 degrees Fahrenheit).
- Some seed types require more extreme treatments to germinate
such as fire. To simulate this, nick the seed coat (with blade
or file) and immerse it in hot water (190 degrees Fahrenheit).
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Asexual Propagation: is
cloning. Offspring are genetically and physically identical to their
parent plant.
Methods:
- Division
- Crown division – the crown is the region where
the roots join the stem(s). In fall, you can cut the crown
longitudinally into several sections, each containing both
stem and root, and plant them immediately or overwinter them
in a root cellar until spring (rhubarb).
- Root division – in fall, after dieback, divide
the root system into chunks, each containing several dormant
underground buds (which will develop into next year's shoots),
and plant immediately. The new plants will require at least
a year to mature.
- Suckers – the shoots that sprout along underground
stems (runners) some distance from original crown will eventually
develop into whole new plant (raspberries).
- Stolons – these above ground runners root periodically,
producing a whole new plant (strawberries).
- Tip rooting – the tip of a stem contacts the
ground and sprouts roots, developing into new plant (blackberries).
- Layering – generating new plants by making plants sprout
roots along the stems.
- Simple layering – this works best with woody
shrubs with flexible stems. In spring, bend the stem to the
ground until it contacts the soil just behind the tip, cover
it with soil behind the tip, and wait a full year before severing
the stem from parent plant. It can help to notch the underside
of the stem where it contacts ground (black raspberries, blueberries,
currants, grapes).
- Compound layering – this is like simple layering,
but cover a trailing stem with soil at several points/nodes
along its length (grapes).
- Mound layering (stooling) – with woody, inflexible/breakable
stems, notch the base of each stem, mound soil around base
of the plant so that notches are well covered, and wait l-2
years before severing the rooted stems from the parent plant
(blueberries, currants, gooseberries, hazelnut shrubs, juneberries).
- Air layering – in mid spring, ring or girdle
a branch in the area of last year's growth, surround it in
a moist substance (such as compost surrounded in moss), wrap
it in plastic, then wait 1 growing season before severing
the branch between parent and plant.
- Cuttings - generating whole new plants from cut pieces of stem
or root:
- Stem cuttings (hardwood) – after fall dormancy
and before spring bud-break, cut 6-12” pieces of the
most recent season's growth, and store at 50-55 degrees Fahrenheit
for 2 months (this simulates the first half of winter) to
induce callus formation (the tissue from which roots can development).
Then store at 35-40 degrees Fahrenheit for 2 months (this
simulates the second half of winter) to slow growth until
spring and facilitate bud break. Pot them in spring, and tent
them in plastic to prevent desiccation until the roots develop
(grapes, currants, filberts, gooseberries, mulberries).
- Root cuttings – in fall after dormancy, cut
roots of pencil thickness into 3-4” sections. Overwinter
them as described for stem cuttings and plant them in spring
(blackberries).
- Grafting – creating a new plant by joining a scion or
bud from a variety with desirable fruit, flowers, leaves, and/or
bark to the roots of a variety with desirable growth characteristics.
This is done with a scion or bud. The graft is successful ('takes')
when the cambium layers/vascular systems of the two varieties
join.
- Scion: a 4-8” stem cutting (2-5 buds) used
to graft. Note: “Water sprouts” are not good scions
for either whip or bud grafting.
- Rootstock: a 2-3 year old seedling, lopped off l-2”
above ground level, affords hardiness to specific soil and
weather conditions, and pest/disease resistance, also influences
the scion's growth potential (dwarfing, in some cases).
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Grafting Techniques:
Simple Whip Grafting
Whip, splice, or cleft grafting is a simple technique for propagating
apples, pears, and to a lesser extent, other fruits and nuts. Several
items are required: rootstock, scionwood (preferably of the same
diameter) sharp grafting knife, rubber strips, and water proof tree
dressing (wax base or tree heal). Timing is important, the ideal
time to graft is mid to rate spring when the sap is flowing well,
apple trees in your area are in full bloom, and night time temperatures
don't go below freezing. The scionwood to be grafted should have
been cut previously and fully dormant. The over-riding principle
of grafting is to match cambium surfaces, that part of the tree
just under the bark that is the tree's bloodstream, try to maximize
the contact of this area between the rootstock and the scionwood.
Before use, scionwood should be refrigerated with moist packing
(paper towels, or newspaper) in a plastic bag.
Step 1
Find the part of the rootstock trunk that is the same diameter is
the scion. Given a choice, partial cambial contact lower on the
stem (say, 4" to 12"), is better than full contact higher
on the stem (say 2' to 3'). Clean off the branchlets in the area.
For a whip graft, use a sharp grafting knife to cut upward through
the rootstock with a sloping, flat cut, Longer is better, about
1 1/2" is usually fine. For a cleft graft, cut across the rootstock,
then split it 1-1½” down, as shown in the figure. Do
not touch the cut surface and work quickly, but safely!
Step 2
Cut your scion to complement the rootstock. For a whip graft, use
a cut identical to the one used on the rootstcok, and of identical
length. For a cleft graft, cut the scion on both sides to make a
wedge about 1¼” long. The scion should have 2 to 3
buds on it. Examine both the scion and the stock next to one another
to be sure they will match. Place the scion on the stock (or, in
the case of cleft grafting, fit it into the split in the stock)
and gently, but firmly, wrap the union with a long rubber strip.
A good guide on tightness is that the rubber strip should never
be stretched out so that it is less than 1/2 the unstretched width.
Tuck the end of the strip under the last wrap. Check again the alignment
of cambial layers.
Step 3
Paint the top cut end of the scion with dressing. Cover the entire
graft union area with dressing to prevent drying out. We use grafting
bags, held at the bottom with rubber strips. This acts like a miniature
greenhouse, callusing more quickly. Removal of this bag should be
in approximately two weeks. Checking first to see if you have good
bud growth. You now have a graft. Care for your new trees following
the procedures in the next section.
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T-Budding
T-budding is a form of grafting useful for all the common fruit
trees and many nuts. It uses the same materials as whip grafting
but at a different time of the year. The best time for budding is
late August in northern areas when the rootstock is still vigorously
growing, but the wood has hardened slightly and the scion buds at
the base of the leaf stems are full sized, mature and plump. Stocks
for budding may be set out the previous spring and cut back to the
ground, to encourage a single vigorous shoot to receive the bud.
Larger trees can also be topworked by budding and buds are always
best placed on the current years growth. Fresh cut scions are important,
as soon as the twigs (use the middle 2/3 of current years growth)
are cut, the leaf blade should be cut off, leaving the leaf stem
as a handle. These scions should be carried with the cut ends in
water, and kept out of the sun, perhaps in a cooler.
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Step 1
Prepare the stock by stripping off the leaves in
the area where the bud will be inserted. Lower is
usually always better, 4” above ground level
on t year whips. Cut into the south side of the
tree with a short horizontal cut, make a downward
vertical cut for 1 1/4” or so, just through
the bark. This is the “T” shape that
gives this the common name of T-budding. Use the
budding nose or bud opener blade on your knife to
lift the 2 side flaps of the bark away from the
wood. |
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| Step 2
Remove the bud from the bud stick by inserting the
knife about 3/4” below the bud and, cutting
just deep enough to remove a sliver of wood below
the bud, draw it under the bud to about 3/4”
above the bud. Remove the bud by its leaf stem,
with a straight cut across the top end of the bud
shield. Keep your fingers and dirt off the cut surfaces! |
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Step 3
Insert the bud shield into the top of the T, opening
the bark flaps of the rootstock, pushing down firmly.
Cut off the upper end of the bud shield, in line
with the horizontal cut, or top of the T, on the
stock. Clean sharp cuts are important in all these
operations. |
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| Step 4
Wrap the bark and bud with a rubber strip, somewhat
more loosely than described in the grafting section.
Do not cover the bud or leaf stem; the latter will
quickly fall off in a week or so, indicating successful
union. In dry or hot conditions, it is useful to
paint on a dressing to prevent drying out. Here
we use saran wrap, to go completely around then
rubber strips both top and bottom, acting like a
miniature green house, callusing more quickly. If
it is not a take, you will find the top or bottom
of the inserted bud starting to pull away (drying
out). The dormant bud stays healed in through the
winter. |
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Step 5
The following spring, just as the buds begin to
shoot, cut off the rootstock l/4” above the
bud union. Use a bamboo stake to tie the new bud
shoot, when it reaches 6” or better in height,
to help in training and prevent dislocation in the
wind. |
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Chip Budding
Chip budding is now becoming the normal procedure used most widely
in the fruit tree propagation field today. This procedure is used
almost exclusively by propagation experts for 2 main reasons, (1)
it is faster and easier to do than T-budding and (2) in times of
variable weather, or plant growth conditions at budding time, it
tends to be more successful than T budding. T-budding is still important
when there is a mismatch in size between budwood and rootstock,
and for the novice (seems more the case as compared to a professionals
view point), therefore, this procedure should still be encouraged
when it “feels” or “looks like” the more
comfortable approach to getting the job done. At Bear Creek we use
both procedures when one or the other budding process looks like
it will work best. The most important facet of chip budding is that
the bud stick matches in caliper the part of the rootstock where
the bud is to be placed. If the match is there between the two,
then you will know you can chip bud as opposed to placing a T-bud.
| Step 1
Prepare the Rootstock as you would for T-budding by stripping
off the leaves in the area where the bud will be inserted. |
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Step 2
Cut the bud chip from your budwood stick, Do this
by inserting the knife 1/2” above the bud
and cutting in a downward stroke under the bud to
a point 1/4" below the bud, but not completely out
of the stock' To remove the bud chip, now take Your
knife and cut under the bud in a slight downward
angle (a dove tail cut), into the stem, matching
up with your first cut. Remove the bud chip. Most
folks hold it between their teeth by the leaf stem
base to prevent contamination. |
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| Step 3
Using the bud stick with the chip removed as a pattern,
(some folks lay it next to the rootstock, others
hold it along side the rootstock between several
fingers), cut a matching chip of wood from the rootstock.
Compare the notch cut on the bud stick to the notch
cut on the rootstock to see that they match. If
your rootstock notch is smaller, (you should always
cut to favor the small size), you can whittle on
it with your knife to size it up to match. |
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Step 4
Insert your bud chip into the notch on the rootstock.
The dovetail section is the most important part
of the “match”, as it creates the solid
seat for the bud chip to sit firmly in place. |
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| Step 5
Wrap the bud chip firmly but not too tight, with
a rubber strip leaving the bud or leaf stem itself
uncovered. Follow up with a “greenhouse wrap”
of saran wrap or parafilm chip budding tape to help
hold in the moisture and warmth to assist the callusing
Process. A good dose of irrigation to provide a
good final boost to the growing process and you
should have a completely “healed in”
bud chip, prepared to go through the winter comfortably.
Parafilm chip budding tape along with the budding
rubber will break down and fall off on their own.
Any other type of wrappings you will need to remove
yourself before winter. |
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| Step 6
The following spring, as you would with a T-budded
rootstock, just as the buds begin to shoot, cut
off the rootstock just above any or all callusing
of the bud union. Also plan to stake the new shoot
when it reaches about 6” high to help train
it and prevent wind breakage. |
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Reprinted and adapted with permission from The Bear Creek Nursery
Catalogue, published by Bear Creek Nursery, which is no longer in
business.
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